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Type	Result	AUM ($B)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
BVF Inc/IL	[2025-06-30]	DFND	0	19,822,436
Wells Fargo & Company/MN	[2025-06-30]	DFND	1	1,024,608
Renaissance Technologies LLC	[2025-06-30]	SOLE	828,321	828,321
Acadian Asset Management LLC	[2025-06-30]	DFND	411,696	411,696
BlackRock, Inc.	[2025-06-30]	SOLE	411,307	411,307
Northern Trust Corp	[2025-06-30]	DFND	381,354	381,354
Millennium Management LLC	[2025-06-30]	SOLE	291,532	291,532
Deutsche Bank AG	[2025-06-30]	DFND	253,375	253,375
Citadel Advisors LLC	[2025-06-30]	DFND	174,499	174,499
Marshall Wace LLP	[2025-06-30]	OTR	0	135,841
Dimensional Fund Advisors LP	[2025-06-30]	SOLE	133,270	133,270
Handelsbanken Fonder AB	[2025-06-30]	SOLE	0	105,000
Eversept Partners LP	[2025-06-30]	SOLE	100,752	100,752
Morgan Stanley	[2025-06-30]	DFND	10,471	72,895
Dimensional Fund Advisors LP	[2025-06-30]	DFND	0	65,416
Geode Capital Management LLC	[2025-06-30]	DFND	63,601	63,601
Two Sigma Investments LP	[2025-06-30]	SOLE	30,690	30,690
Cantor Fitzgerald L P	[2025-06-30]	DFND	0	30,000
Two Sigma Advisers LP	[2025-06-30]	SOLE	27,010	28,810
SEI Investments Co	[2025-06-30]	SOLE	24,897	24,897
Banque Cantonale Vaudoise	[2025-06-30]	SOLE	18,440	18,440
Squarepoint Ops LLC	[2025-06-30]	DFND	0	12,642
Goldman Sachs Group Inc	[2025-06-30]	DFND	10,369	10,369
Quinn Opportunity Partners LLC	[2025-06-30]	SOLE	10,100	10,100
Two Sigma Securities LLC	[2025-06-30]	SOLE	10,085	10,085
Silverberg Bernstein Capital Management LLC	[2025-06-30]	SOLE	0	10,000
Barclays PLC	[2025-06-30]	SOLE	7,907	7,907
UBS Group AG	[2025-06-30]	DFND	1,466	7,467
JPMorgan Chase & Co	[2025-06-30]	DFND	6,110	6,110
Bank of America Corp /DE/	[2025-06-30]	DFND	0	5,573
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