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Type	Result	AUM ($B)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
FMR LLC	[2023-03-31]	DFND	12,979,080.0	12,979,080.0
Pentwater Capital Management LP	[2023-03-31]	SOLE	5,975,000.0	5,975,000.0
SilverArc Capital Management LLC	[2023-03-31]	SOLE	2,689,301.0	2,689,301.0
EcoR1 Capital LLC	[2023-03-31]	SOLE	2,650,000.0	2,650,000.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	2,302,145.0
BlackRock Inc.	[2023-03-31]	SOLE	1,868,199.0	1,889,823.0
Paradigm Biocapital Advisors LP	[2023-03-31]	SOLE	1,072,410.0	1,072,410.0
Aisling Capital Management LP	[2023-03-31]	SOLE	1,043,267.0	1,043,267.0
Acadian Asset Management LLC	[2023-03-31]	DFND	1,011,383.0	1,011,383.0
Cormorant Asset Management LP	[2023-03-31]	DFND	1,000,000.0	1,000,000.0
Millennium Management LLC	[2023-03-31]	SOLE	975,612.0	975,612.0
Dimensional Fund Advisors LP	[2023-03-31]	SOLE	672,877.0	695,846.0
Susquehanna International Group LLP	[2023-03-31]	DFND	651,394.0	651,394.0
Renaissance Technologies LLC	[2023-03-31]	SOLE	341,031.0	603,200.0
Laurion Capital Management LP	[2023-03-31]	SOLE	545,700.0	545,700.0
Geode Capital Management LLC	[2023-03-31]	DFND	506,090.0	506,090.0
Morgan Stanley	[2023-03-31]	DFND	475,806.0	481,826.0
Two Sigma Investments LP	[2023-03-31]	SOLE	481,044.0	481,044.0
Marshall Wace LLP	[2023-03-31]	OTR	0.0	448,745.0
Wellington Management Group LLP	[2023-03-31]	DFND	0.0	372,285.0
Goldman Sachs Group Inc	[2023-03-31]	DFND	293,362.0	293,362.0
Cubist Systematic Strategies, LLC	[2023-03-31]	DFND	0.0	292,721.0
Qube Research & Technologies Ltd	[2023-03-31]	SOLE	0.0	277,168.0
Citadel Advisors LLC	[2023-03-31]	DFND	274,704.0	274,704.0
Charles Schwab Investment Management Inc	[2023-03-31]	SOLE	260,746.0	260,746.0
Advisory Services Network LLC	[2023-03-31]	SOLE	257,271.0	257,271.0
Jane Street Group LLC	[2023-03-31]	DFND	253,169.0	253,169.0
Bank of America Corp /DE/	[2023-03-31]	DFND	0.0	252,148.0
State Street Corp	[2023-03-31]	DFND	234,467.0	234,467.0
HRT Financial LP	[2023-03-31]	DFND	191,064.0	191,064.0
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