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Type	Result	AUM ($B)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
Ameriprise Financial Inc	[2025-06-30]	DFND	0	4,236,877
Portolan Capital Management LLC	[2025-06-30]	SOLE	3,633,622	3,633,622
BlackRock, Inc.	[2025-06-30]	SOLE	2,848,745	2,924,699
Vanguard Group Inc	[2025-06-30]	SOLE	0	2,500,667
Kent Lake PR LLC	[2025-06-30]	SOLE	2,301,000	2,301,000
Invenomic Capital Management LP	[2025-06-30]	SOLE	1,144,904	1,144,904
William Blair Investment Management LLC	[2025-06-30]	SOLE	911,671	1,127,981
Easterly Investment Partners LLC	[2025-06-30]	OTR	0	1,083,635
Blue Water Life Science Advisors LP	[2025-06-30]	SOLE	957,747	957,747
Geode Capital Management LLC	[2025-06-30]	DFND	887,080	887,080
Morgan Stanley	[2025-06-30]	DFND	762,847	771,440
Balyasny Asset Management LP	[2025-06-30]	SOLE	767,801	767,801
State Street Corp	[2025-06-30]	DFND	710,317	764,855
Dimensional Fund Advisors LP	[2025-06-30]	SOLE	685,857	726,596
Williams Jones Wealth Management LLC	[2025-06-30]	SOLE	0	676,014
Trium Capital LLP	[2025-06-30]	SOLE	586,425	586,425
Tikvah Management LLC	[2025-06-30]	SOLE	571,619	571,619
Parkman Healthcare Partners LLC	[2025-06-30]	SOLE	515,992	515,992
Marshall Wace LLP	[2025-06-30]	OTR	0	444,191
Russell Investments Group, Ltd.	[2025-06-30]	OTR	0	424,042
Aberdeen Group plc	[2025-06-30]	DFND	420,856	420,856
Baillie Gifford & Co	[2025-06-30]	DFND	391,135	391,135
UBS Group AG	[2025-06-30]	DFND	384,626	384,951
SCS Capital Management LLC	[2025-06-30]	SOLE	354,339	354,339
Schonfeld Strategic Advisors LLC	[2025-06-30]	SOLE	325,677	325,677
Northern Trust Corp	[2025-06-30]	DFND	315,442	315,442
Charles Schwab Investment Management Inc	[2025-06-30]	SOLE	298,989	298,989
State of Wisconsin Investment Board	[2025-06-30]	SOLE	289,759	289,759
Citigroup Inc	[2025-06-30]	DFND	280,578	280,578
JPMorgan Chase & Co	[2025-06-30]	DFND	243,101	260,501
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