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Type	Result	AUM ($B)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
SR One Capital Management LP	[2023-03-31]	SOLE	6,822,114.0	6,822,114.0
Carlyle Group Inc.	[2023-03-31]	DFND	0.0	4,822,114.0
Column Group LLC	[2023-03-31]	SOLE	4,309,329.0	4,309,329.0
Sectoral Asset Management Inc	[2023-03-31]	SOLE	2,382,644.0	2,382,644.0
Parallel Advisors LLC	[2023-03-31]	SOLE	1,693,559.0	1,693,559.0
Blackstone Inc.	[2023-03-31]	OTR	0.0	1,308,587.0
OUP Management Co LLC	[2023-03-31]	SOLE	1,025,697.0	1,025,697.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	405,919.0
Sea Otter Advisors LLC	[2023-03-31]	SOLE	263,942.0	263,942.0
Renaissance Technologies LLC	[2023-03-31]	SOLE	7,000.0	143,900.0
Geode Capital Management LLC	[2023-03-31]	DFND	104,982.0	104,982.0
Oasis Management Co Ltd.	[2023-03-31]	SOLE	100,000.0	100,000.0
Susquehanna International Group LLP	[2023-03-31]	DFND	62,629.0	62,629.0
BlackRock Inc.	[2023-03-31]	SOLE	53,076.0	53,076.0
Wescott Financial Advisory Group LLC	[2023-03-31]	SOLE	0.0	30,556.0
Northern Trust Corp	[2023-03-31]	DFND	9,248.0	25,993.0
Two Sigma Securities LLC	[2023-03-31]	SOLE	24,635.0	24,635.0
UBS Group AG	[2023-03-31]	SOLE	21,414.0	21,414.0
State Street Corp	[2023-03-31]	DFND	20,085.0	20,085.0
MGO One Seven LLC	[2023-03-31]	SOLE	20,070.0	20,070.0
Callan Capital LLC	[2023-03-31]	SOLE	0.0	20,000.0
EP Wealth Advisors LLC	[2023-03-31]	SOLE	6,256.0	17,607.0
Ground Swell Capital LLC	[2023-03-31]	SOLE	11,752.0	11,752.0
Tower Research Capital LLC (TRC)	[2023-03-31]	DFND	10,316.0	10,316.0
Advisor Group Holdings Inc	[2023-03-31]	DFND	1,500.0	1,500.0
Bank of America Corp /DE/	[2023-03-31]	DFND	0.0	100.0
Wells Fargo & Company/MN	[2023-03-31]	DFND	75.0	75.0
Concourse Financial Group Securities Inc	[2023-03-31]	SOLE	0.0	0.0
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