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Type	Result	AUM ($BB)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
Flagship Pioneering Inc	[2023-03-31]	DFND	0.0	12,674,120.0
FMR LLC	[2023-03-31]	DFND	3,014,271.0	3,015,012.0
Artal Group S.A.	[2023-03-31]	DFND	1,720,720.0	1,720,720.0
Euclidean Capital LLC	[2023-03-31]	SOLE	1,261,261.0	1,261,261.0
BlackRock Inc.	[2023-03-31]	SOLE	1,223,927.0	1,242,520.0
Raymond James & Associates	[2023-03-31]	SOLE	0.0	1,078,056.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	984,935.0
Deerfield Management Company LP Series C	[2023-03-31]	DFND	777,525.0	777,525.0
Alphabet Inc.	[2023-03-31]	DFND	0.0	500,901.0
State Street Corp	[2023-03-31]	DFND	385,791.0	409,513.0
Geode Capital Management LLC	[2023-03-31]	DFND	386,008.0	386,008.0
Driehaus Capital Management LLC	[2023-03-31]	SOLE	343,720.0	343,720.0
Birchview Capital LP	[2023-03-31]	SOLE	264,843.0	264,843.0
Northern Trust Corp	[2023-03-31]	DFND	4,877.0	186,075.0
Citadel Advisors LLC	[2023-03-31]	DFND	156,372.0	156,372.0
Affinity Asset Advisors LLC	[2023-03-31]	SOLE	136,771.0	136,771.0
Jacobs Levy Equity Management Inc	[2023-03-31]	SOLE	130,847.0	130,847.0
Morgan Stanley	[2023-03-31]	DFND	18,229.0	93,567.0
Bank of New York Mellon Corp	[2023-03-31]	DFND	68,271.0	71,032.0
Charles Schwab Investment Management Inc	[2023-03-31]	SOLE	54,929.0	54,929.0
Nuveen Asset Management LLC	[2023-03-31]	DFND	53,480.0	53,480.0
Baker Bros Advisors LP	[2023-03-31]	SOLE	45,027.0	45,027.0
Millennium Management LLC	[2023-03-31]	SOLE	37,885.0	37,885.0
Vanguard Group Inc	[2023-03-31]	DFND	0.0	37,030.0
Man Group plc	[2023-03-31]	DFND	33,977.0	33,977.0
Goldman Sachs Group Inc	[2023-03-31]	DFND	31,065.0	31,075.0
Russell Investments Group, Ltd.	[2023-03-31]	DFND	0.0	30,017.0
Acadian Asset Management LLC	[2023-03-31]	DFND	27,085.0	27,085.0
Cornercap Investment Counsel Inc	[2023-03-31]	SOLE	17,722.0	26,163.0
RhumbLine Advisers	[2023-03-31]	SOLE	9,805.0	25,314.0
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