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Type	Result	AUM ($B)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
Blackstone Inc	[2021-12-31]	OTR	0.0	3,375,037.0
Omega Fund Management (US) Inc.	[2021-12-31]	SOLE	3,018,737.0	3,018,737.0
Frazier Management LLC	[2021-12-31]	SOLE	2,971,274.0	2,971,274.0
FMR LLC	[2021-12-31]	DFND	441,177.0	2,849,595.0
Farallon Capital Management LLC	[2021-12-31]	OTR	2,471,313.0	2,471,313.0
Citadel Advisors LLC	[2021-12-31]	DFND	2,452,926.0	2,452,926.0
Price T Rowe Associates Inc /MD/	[2021-12-31]	SOLE	308,289.0	2,241,917.0
BlackRock Inc.	[2021-12-31]	SOLE	1,332,133.0	1,336,782.0
Woodline Partners LP	[2021-12-31]	SOLE	848,129.0	848,129.0
Emerald Advisers Inc/Pa	[2021-12-31]	SOLE	376,285.0	576,159.0
Emerald Mutual Fund Advisers Trust	[2021-12-31]	SOLE	540,364.0	540,364.0
StepStone Group LP	[2021-12-31]	DFND	522,987.0	522,987.0
Vanguard Group Inc	[2021-12-31]	SOLE	0.0	446,587.0
Driehaus Capital Management LLC	[2021-12-31]	SOLE	299,548.0	361,819.0
Adage Capital Partners GP LLC	[2021-12-31]	SOLE	275,000.0	275,000.0
Kingdon Capital Management LLC	[2021-12-31]	DFND	173,830.0	173,830.0
Geode Capital Management LLC	[2021-12-31]	DFND	164,613.0	164,613.0
Millennium Management LLC	[2021-12-31]	SOLE	151,786.0	151,786.0
State Street Corp	[2021-12-31]	DFND	142,036.0	150,791.0
Parkman Healthcare Partners LLC	[2021-12-31]	SOLE	110,000.0	110,000.0
Northern Trust Corp	[2021-12-31]	DFND	1,711.0	103,809.0
Goldman Sachs Group Inc	[2021-12-31]	DFND	87,757.0	87,762.0
HRT Financial LLC	[2021-12-31]	DFND	65,989.0	65,989.0
Kennedy Capital Management Inc	[2021-12-31]	SOLE	53,903.0	53,903.0
Alps Advisors Inc	[2021-12-31]	DFND	38,802.0	38,802.0
D. E. Shaw & Co., Inc.	[2021-12-31]	DFND	32,058.0	32,058.0
Susquehanna International Group LLP	[2021-12-31]	DFND	28,274.0	28,274.0
Bank of New York Mellon Corp	[2021-12-31]	DFND	21,932.0	27,968.0
Swiss National Bank	[2021-12-31]	SOLE	25,800.0	25,800.0
Charles Schwab Investment Management Inc	[2021-12-31]	SOLE	25,188.0	25,188.0
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