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Type	Result	AUM ($BB)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
Granahan Investment Management LLC	[2023-03-31]	SOLE	1,056,831.0	1,282,710.0
FMR LLC	[2023-03-31]	DFND	1,046,117.0	1,047,033.0
Hood River Capital Management LLC	[2023-03-31]	SOLE	965,951.0	1,044,929.0
BlackRock Inc.	[2023-03-31]	SOLE	971,554.0	985,397.0
Wasatch Advisors LP	[2023-03-31]	SOLE	923,928.0	953,318.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	721,609.0
Marshall Wace LLP	[2023-03-31]	OTR	0.0	681,222.0
Janus Henderson Group PLC	[2023-03-31]	DFND	0.0	652,672.0
Federated Hermes Inc	[2023-03-31]	DFND	625,000.0	625,000.0
FIL Ltd	[2023-03-31]	DFND	595,582.0	595,582.0
D. E. Shaw & Co., Inc.	[2023-03-31]	DFND	562,316.0	562,316.0
Millennium Management LLC	[2023-03-31]	SOLE	526,342.0	526,342.0
AWM Investment Company Inc	[2023-03-31]	SOLE	452,096.0	452,096.0
Renaissance Technologies LLC	[2023-03-31]	SOLE	449,700.0	449,700.0
Citadel Advisors LLC	[2023-03-31]	DFND	407,004.0	407,004.0
Dimensional Fund Advisors LP	[2023-03-31]	SOLE	371,328.0	389,600.0
Nuveen Asset Management LLC	[2023-03-31]	DFND	372,007.0	372,007.0
Kornitzer Capital Management Inc /KS	[2023-03-31]	SOLE	281,799.0	281,799.0
Geode Capital Management LLC	[2023-03-31]	DFND	280,577.0	280,577.0
State Street Corp	[2023-03-31]	DFND	235,464.0	266,008.0
Bank of New York Mellon Corp	[2023-03-31]	DFND	171,059.0	205,328.0
Cubist Systematic Strategies, LLC	[2023-03-31]	DFND	0.0	179,786.0
Morgan Stanley	[2023-03-31]	DFND	166,703.0	177,347.0
Balyasny Asset Management LP	[2023-03-31]	SOLE	177,035.0	177,035.0
Skylands Capital LLC	[2023-03-31]	SOLE	175,515.0	175,515.0
Ameriprise Financial Inc	[2023-03-31]	DFND	0.0	171,649.0
Two Sigma Investments LP	[2023-03-31]	SOLE	167,099.0	167,099.0
Voloridge Investment Management LLC	[2023-03-31]	SOLE	138,651.0	138,651.0
Bank of New York Mellon Corp	[2023-03-31]	OTR	129,803.0	129,803.0
Northern Trust Corp	[2023-03-31]	DFND	8,205.0	129,541.0
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