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Type	Result	AUM ($B)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
Perceptive Advisors LLC	[2025-06-30]	SOLE	5,402,964	5,402,964
BML Capital Management LLC	[2025-06-30]	SOLE	0	3,230,667
Vanguard Group Inc	[2025-06-30]	SOLE	0	1,524,656
Propel Bio Management LLC	[2025-06-30]	DFND	0	1,492,792
Simplify Asset Management Inc	[2025-06-30]	SOLE	1,492,792	1,492,792
Tang Capital Management LLC	[2025-06-30]	OTR	0	600,000
Bank of America Corp /DE/	[2025-06-30]	DFND	415,000	416,203
Acadian Asset Management LLC	[2025-06-30]	DFND	405,481	405,481
BlackRock, Inc.	[2025-06-30]	SOLE	376,454	376,454
Pathstone Holdings, LLC	[2025-06-30]	SOLE	0	372,196
Geode Capital Management LLC	[2025-06-30]	DFND	305,964	305,964
Renaissance Technologies LLC	[2025-06-30]	SOLE	241,000	241,000
Citadel Advisors LLC	[2025-06-30]	DFND	234,228	234,228
UBS Group AG	[2025-06-30]	DFND	202,543	203,475
Qube Research & Technologies Ltd	[2025-06-30]	SOLE	192,684	192,684
Two Sigma Advisers LP	[2025-06-30]	SOLE	143,100	154,200
Laird Norton Wetherby Trust Company, LLC	[2025-06-30]	SOLE	127,309	127,309
State Street Corp	[2025-06-30]	DFND	107,381	107,381
Susquehanna International Group LLP	[2025-06-30]	OTR	80,524	80,524
Osaic Holdings Inc	[2025-06-30]	DFND	0	63,500
Northern Trust Corp	[2025-06-30]	DFND	53,609	53,609
Clarius Group, LLC	[2025-06-30]	SOLE	53,551	53,551
XTX Topco Ltd	[2025-06-30]	DFND	52,986	52,986
Integrity Financial Corp /WA	[2025-06-30]	SOLE	0	52,902
	[2025-06-30]	SOLE	0	35,765
Brighton Jones LLC	[2025-06-30]	SOLE	0	32,200
Royal Bank of Canada	[2025-06-30]	DFND	27,156	27,156
Virtu Financial LLC	[2025-06-30]	DFND	24,742	24,742
Simon Quick Advisors LLC	[2025-06-30]	SOLE	0	20,538
Journey Strategic Wealth LLC	[2025-06-30]	SOLE	0	20,538
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