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Type	Result	AUM ($B)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
Lynx1 Capital Management LP	[2025-06-30]	OTR	0	1,538,573
Ikarian Capital LLC	[2025-06-30]	SOLE	599,913	599,913
Opaleye Management Inc	[2025-06-30]	SOLE	375,000	375,000
Boothbay Fund Management LLC	[2025-06-30]	OTR	0	159,448
Vanguard Group Inc	[2025-06-30]	SOLE	0	138,401
Geode Capital Management LLC	[2025-06-30]	DFND	112,301	112,301
Adar1 Capital Management LLC	[2025-06-30]	SOLE	54,819	54,819
BlackRock, Inc.	[2025-06-30]	SOLE	42,727	42,727
Virtu Financial LLC	[2025-06-30]	DFND	40,181	40,181
State Street Corp	[2025-06-30]	DFND	33,699	33,699
UBS Group AG	[2025-06-30]	DFND	28,602	28,603
Two Sigma Securities LLC	[2025-06-30]	SOLE	28,562	28,562
Northern Trust Corp	[2025-06-30]	DFND	23,044	23,044
Dauntless Investment Group, LLC	[2025-06-30]	SOLE	0	20,000
Vanguard Group Inc	[2025-06-30]	DFND	0	5,460
Tower Research Capital LLC (TRC)	[2025-06-30]	DFND	2,107	2,107
Osaic Holdings Inc	[2025-06-30]	DFND	0	353
SBI Securities Co., Ltd.	[2025-06-30]	DFND	223	223
Bank of America Corp /DE/	[2025-06-30]	DFND	0	157
Arax Advisory Partners	[2025-06-30]	SOLE	0	91
Morgan Stanley	[2025-06-30]	DFND	42	49
Atlantic Union Bankshares Corp	[2025-06-30]	SOLE	0	14
SOA Wealth Advisors LLC	[2025-06-30]	SOLE	13	13
BNP Paribas Financial Markets	[2025-06-30]	DFND	0	5
Activest Wealth Management	[2025-06-30]	SOLE	0	4
City Holding Co	[2025-06-30]	SOLE	4	4
Financial Gravity Companies, Inc.	[2025-06-30]	SOLE	2	2
JPMorgan Chase & Co	[2025-06-30]	DFND	2	2
Steward Partners Investment Advisory LLC	[2025-06-30]	SOLE	0	1
FMR LLC	[2025-06-30]	DFND	0	1
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