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Type	Result	AUM ($BB)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
KCK Ltd	[2023-03-31]	SOLE	5,621,111.0	5,621,111.0
OrbiMed Advisors LLC	[2023-03-31]	SOLE	4,012,281.0	4,012,281.0
Soleus Capital Management LP	[2023-03-31]	SOLE	2,428,714.0	2,428,714.0
Morgan Stanley	[2023-03-31]	DFND	2,256,291.0	2,258,016.0
FMR LLC	[2023-03-31]	DFND	1,492,931.0	1,492,931.0
Millennium Management LLC	[2023-03-31]	SOLE	731,610.0	731,610.0
Nantahala Capital Management LLC	[2023-03-31]	SOLE	425,925.0	425,925.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	294,041.0
Kent Lake Capital LLC	[2023-03-31]	SOLE	127,676.0	127,676.0
Geode Capital Management LLC	[2023-03-31]	DFND	71,515.0	71,515.0
Bridgeway Capital Management LLC	[2023-03-31]	SOLE	51,648.0	51,648.0
BlackRock Inc.	[2023-03-31]	SOLE	32,471.0	32,471.0
SG Americas Securities LLC	[2023-03-31]	SOLE	25,388.0	25,388.0
State Street Corp	[2023-03-31]	DFND	22,451.0	22,451.0
Northern Trust Corp	[2023-03-31]	DFND	0.0	16,171.0
Renaissance Technologies LLC	[2023-03-31]	SOLE	14,500.0	14,500.0
Susquehanna International Group LLP	[2023-03-31]	DFND	14,052.0	14,052.0
Bank of America Corp /DE/	[2023-03-31]	DFND	0.0	13,016.0
Financial Advocates Investment Management	[2023-03-31]	SOLE	0.0	10,000.0
UBS Group AG	[2023-03-31]	SOLE	2,623.0	2,623.0
Vanguard Group Inc	[2023-03-31]	DFND	0.0	2,507.0
Tower Research Capital LLC (TRC)	[2023-03-31]	DFND	784.0	784.0
Bergankdv Wealth Management LLC	[2023-03-31]	SOLE	420.0	420.0
Wells Fargo & Company/MN	[2023-03-31]	DFND	69.0	219.0
Barclays PLC	[2023-03-31]	SOLE	118.0	118.0
Concourse Financial Group Securities Inc	[2023-03-31]	SOLE	0.0	0.0
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