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Type	Result	AUM ($B)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
Vanguard Group Inc	[2025-06-30]	SOLE	0	3,851,671
BlackRock, Inc.	[2025-06-30]	SOLE	3,649,100	3,834,604
BVF Inc/IL	[2025-06-30]	DFND	0	1,690,221
RTW Investments LP	[2025-06-30]	SOLE	1,516,903	1,516,903
Macquarie Group Ltd	[2025-06-30]	SOLE	1,113,166	1,137,748
Geode Capital Management LLC	[2025-06-30]	DFND	989,453	989,453
FMR LLC	[2025-06-30]	DFND	974,852	976,730
State Street Corp	[2025-06-30]	DFND	845,069	958,503
Fairmount Funds Management LLC	[2025-06-30]	SOLE	815,176	815,176
Deep Track Capital LP	[2025-06-30]	SOLE	801,371	801,371
Bank of America Corp /DE/	[2025-06-30]	DFND	27,724	753,429
Eventide Asset Management LLC	[2025-06-30]	SOLE	725,672	725,672
Wellington Management Group LLP	[2025-06-30]	DFND	0	725,044
Alethea Capital Management LLC	[2025-06-30]	SOLE	667,368	667,368
UBS Group AG	[2025-06-30]	DFND	585,093	595,455
Susquehanna International Group LLP	[2025-06-30]	OTR	106,021	576,421
Invesco Ltd.	[2025-06-30]	DFND	0	571,549
Man Group plc	[2025-06-30]	DFND	525,069	525,069
PFM Health Sciences LP	[2025-06-30]	DFND	0	483,105
Norges Bank	[2025-06-30]	SOLE	468,329	468,329
Morgan Stanley	[2025-06-30]	DFND	200,922	447,919
Adage Capital Partners GP LLC	[2025-06-30]	SOLE	447,000	447,000
OrbiMed Advisors LLC	[2025-06-30]	DFND	38,500	427,900
Pictet Asset Management Holding SA	[2025-06-30]	SOLE	372,097	411,850
Northern Trust Corp	[2025-06-30]	DFND	394,099	395,055
Qube Research & Technologies Ltd	[2025-06-30]	SOLE	391,617	391,617
Hood River Capital Management LLC	[2025-06-30]	SOLE	360,034	371,104
Point72 Asset Management LP	[2025-06-30]	DFND	0	369,473
Charles Schwab Investment Management Inc	[2025-06-30]	SOLE	356,651	356,651
Principal Financial Group Inc	[2025-06-30]	DFND	355,511	355,511
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