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Type	Result	AUM ($B)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
FMR LLC	[2020-09-30]	DFND	2.8	10.3
Wellington Management Co LLP	[2020-09-30]	DFND	0.0	6.7
RA Capital Management LP	[2020-09-30]	SOLE	4.8	4.8
BlackRock Inc.	[2020-09-30]	SOLE	4.4	4.5
Vanguard Group Inc	[2020-09-30]	SOLE	0.0	4.4
State Street Corp	[2020-09-30]	DFND	1.8	2.0
Cormorant Asset Management LP	[2020-09-30]	DFND	1.4	1.6
Pictet Asset Management Ltd	[2020-09-30]	DFND	0.0	1.3
Wasatch Advisors Inc	[2020-09-30]	SOLE	0.9	1.1
Marshall Wace North America LP	[2020-09-30]	OTR	0.0	0.9
Baker Bros Advisors LLC	[2020-09-30]	SOLE	0.9	0.9
Macquarie Group Ltd	[2020-09-30]	SOLE	0.9	0.9
Polar Capital LLP	[2020-09-30]	SOLE	0.8	0.8
D. E. Shaw & Co., Inc.	[2020-09-30]	DFND	0.8	0.8
Geode Capital Management LLC	[2020-09-30]	DFND	0.7	0.7
Invesco Ltd.	[2020-09-30]	DFND	0.0	0.7
Point72 Asset Management LP	[2020-09-30]	DFND	0.0	0.7
Dimensional Fund Advisors LP	[2020-09-30]	DFND	0.6	0.6
Northern Trust Corp	[2020-09-30]	DFND	0.1	0.6
Lord Abbett & Co LLC	[2020-09-30]	SOLE	0.5	0.5
Citadel Advisors LLC	[2020-09-30]	DFND	0.5	0.5
Nuveen Asset Management LLC	[2020-09-30]	DFND	0.5	0.5
Goldman Sachs Group Inc	[2020-09-30]	DFND	0.5	0.5
Loomis Sayles & Co L P	[2020-09-30]	DFND	0.5	0.5
Susquehanna International Group LLP	[2020-09-30]	DFND	0.0	0.4
Emerald Advisers LLC	[2020-09-30]	SOLE	0.2	0.4
Emerald Mutual Fund Advisers Trust	[2020-09-30]	SOLE	0.4	0.4
Charles Schwab Investment Management Inc	[2020-09-30]	SOLE	0.4	0.4
Opaleye Management Inc	[2020-09-30]	SOLE	0.4	0.4
Eagle Asset Management Inc	[2020-09-30]	SOLE	0.4	0.4
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