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Type	Result	AUM ($B)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
Opaleye Management Inc	[2025-06-30]	SOLE	2,860,000	2,860,000
EcoR1 Capital LLC	[2025-06-30]	SOLE	2,030,544	2,030,544
BlackRock, Inc.	[2025-06-30]	SOLE	1,501,534	1,531,711
Vanguard Group Inc	[2025-06-30]	SOLE	0	1,278,208
Nantahala Capital Management LLC	[2025-06-30]	SOLE	1,175,951	1,175,951
Geode Capital Management LLC	[2025-06-30]	DFND	592,865	592,865
Morgan Stanley	[2025-06-30]	DFND	477,215	509,568
Divisadero Street Capital Management LP	[2025-06-30]	SOLE	497,716	497,716
Westside Investment Management, Inc.	[2025-06-30]	SOLE	0	472,790
Renaissance Technologies LLC	[2025-06-30]	SOLE	405,900	405,900
Cannell Capital LLC	[2025-06-30]	SOLE	388,175	388,175
State Street Corp	[2025-06-30]	DFND	287,348	323,194
Soleus Capital Management LP	[2025-06-30]	SOLE	285,000	285,000
Northern Trust Corp	[2025-06-30]	DFND	258,272	258,272
JPMorgan Chase & Co	[2025-06-30]	DFND	230,465	242,580
Susquehanna International Group LLP	[2025-06-30]	OTR	30,598	240,898
Connor Clark & Lunn Investment Management Ltd	[2025-06-30]	SOLE	222,083	222,083
Two Sigma Investments LP	[2025-06-30]	SOLE	214,552	214,552
UBS Group AG	[2025-06-30]	DFND	212,484	212,484
Aristides Capital LLC	[2025-06-30]	SOLE	167,587	167,587
Bridgeway Capital Management LLC	[2025-06-30]	SOLE	135,270	135,270
Bank of New York Mellon Corp	[2025-06-30]	DFND	126,350	127,344
Goldman Sachs Group Inc	[2025-06-30]	DFND	109,369	109,379
Stifel Financial Corp	[2025-06-30]	DFND	106,846	108,168
APIS Capital Advisors LLC	[2025-06-30]	SOLE	106,000	106,000
Two Sigma Advisers LP	[2025-06-30]	SOLE	98,400	100,900
Squarepoint Ops LLC	[2025-06-30]	DFND	0	93,796
Raymond James Financial Inc	[2025-06-30]	DFND	12,621	90,415
BNP Paribas Financial Markets	[2025-06-30]	DFND	0	87,696
Anchor Capital Advisors LLC	[2025-06-30]	SOLE	86,617	87,661
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