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Type	Result	AUM ($BB)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
BlackRock Inc.	[2023-03-31]	SOLE	9,881,889.0	10,139,020.0
State Street Corp	[2023-03-31]	DFND	5,390,417.0	5,496,253.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	4,340,909.0
FMR LLC	[2023-03-31]	DFND	3,309,496.0	3,309,836.0
Octagon Capital Advisors LP	[2023-03-31]	SOLE	2,172,000.0	2,172,000.0
Citadel Advisors LLC	[2023-03-31]	DFND	1,838,908.0	1,838,908.0
Invus Financial Advisors LLC	[2023-03-31]	SOLE	1,612,077.0	1,612,077.0
PFM Health Sciences LP	[2023-03-31]	DFND	0.0	1,559,947.0
Boxer Capital, LLC	[2023-03-31]	OTR	1,432,200.0	1,432,200.0
Pictet Asset Management Sa	[2023-03-31]	DFND	1.0	1,283,843.0
Dimensional Fund Advisors LP	[2023-03-31]	SOLE	1,138,657.0	1,172,644.0
Bank of America Corp /DE/	[2023-03-31]	DFND	11,824.0	1,149,218.0
AllianceBernstein LP	[2023-03-31]	DFND	910,823.0	1,017,929.0
Geode Capital Management LLC	[2023-03-31]	DFND	978,167.0	978,167.0
Decheng Capital Management III (Cayman), LLC	[2023-03-31]	SOLE	15,864,970.0	869,790.0
Goldman Sachs Group Inc	[2023-03-31]	DFND	779,781.0	804,534.0
BNP Paribas Asset Management Holding S.A.	[2023-03-31]	DFND	0.0	774,792.0
Granahan Investment Management LLC	[2023-03-31]	SOLE	449,864.0	576,748.0
Morgan Stanley	[2023-03-31]	DFND	518,087.0	556,081.0
JPMorgan Chase & Co	[2023-03-31]	DFND	400,273.0	539,995.0
Boothbay Fund Management LLC	[2023-03-31]	OTR	0.0	532,381.0
Northern Trust Corp	[2023-03-31]	DFND	11,580.0	529,789.0
Susquehanna International Group LLP	[2023-03-31]	DFND	262,120.0	519,020.0
Bank of New York Mellon Corp	[2023-03-31]	DFND	421,653.0	457,210.0
Nuveen Asset Management LLC	[2023-03-31]	DFND	441,241.0	441,241.0
ExodusPoint Capital Management LP	[2023-03-31]	OTR	0.0	427,212.0
Charles Schwab Investment Management Inc	[2023-03-31]	SOLE	421,691.0	421,691.0
Walleye Capital LLC	[2023-03-31]	SOLE	420,087.0	420,087.0
Rafferty Asset Management LLC	[2023-03-31]	SOLE	410,992.0	410,992.0
UBS Asset Management Americas Inc	[2023-03-31]	SOLE	378,363.0	392,323.0
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