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Type	Result	AUM ($BB)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
OrbiMed Advisors LLC	[2023-03-31]	SOLE	73,114,592.0	73,114,592.0
Altium Capital Management LP	[2023-03-31]	SOLE	6,227,559.0	6,227,559.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	1,279,052.0
Renaissance Technologies LLC	[2023-03-31]	SOLE	602,810.0	602,810.0
Crescent Grove Advisors LLC	[2023-03-31]	SOLE	0.0	262,024.0
Bridgeway Capital Management LLC	[2023-03-31]	SOLE	114,500.0	114,500.0
Virtu Financial LLC	[2023-03-31]	DFND	75,240.0	75,240.0
Midwest Professional Planners Ltd	[2023-03-31]	SOLE	0.0	40,744.0
State Street Corp	[2023-03-31]	DFND	23,666.0	23,666.0
Geode Capital Management LLC	[2023-03-31]	DFND	23,593.0	23,593.0
Citadel Advisors LLC	[2023-03-31]	DFND	22,503.0	22,503.0
Northern Trust Corp	[2023-03-31]	DFND	0.0	21,061.0
Resources Investment Advisors LLC	[2023-03-31]	SOLE	0.0	17,255.0
Two Sigma Securities LLC	[2023-03-31]	SOLE	16,120.0	16,120.0
UBS Group AG	[2023-03-31]	SOLE	7,063.0	7,063.0
Tower Research Capital LLC (TRC)	[2023-03-31]	DFND	5,279.0	5,279.0
Morgan Stanley	[2023-03-31]	DFND	0.0	311.0
Advisor Group Holdings Inc	[2023-03-31]	SOLE	0.0	291.0
Wells Fargo & Company/MN	[2023-03-31]	DFND	195.0	195.0
BlackRock Inc.	[2023-03-31]	SOLE	143.0	143.0
FMR LLC	[2023-03-31]	DFND	0.0	88.0
Advisor Group Holdings Inc	[2023-03-31]	DFND	25.0	25.0
Concourse Financial Group Securities Inc	[2023-03-31]	SOLE	0.0	0.0
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