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Type	Result	AUM ($B)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
FMR LLC	[2023-03-31]	DFND	14,854,006.0	14,855,478.0
Great Point Partners LLC	[2023-03-31]	DFND	0.0	7,894,736.0
Bleichroeder LP	[2023-03-31]	SOLE	4,045,132.0	4,045,132.0
Kingdon Capital Management LLC	[2023-03-31]	DFND	3,649,237.0	3,649,237.0
Velan Capital Investment Management LP	[2023-03-31]	SOLE	3,496,861.0	3,496,861.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	3,119,446.0
Massachusetts Financial Services Co /MA/	[2023-03-31]	SOLE	2,874,930.0	2,874,930.0
Nantahala Capital Management LLC	[2023-03-31]	SOLE	1,450,000.0	1,450,000.0
Rice Hall James & Associates LLC	[2023-03-31]	SOLE	605,569.0	771,960.0
Easterly Investment Partners LLC	[2023-03-31]	SOLE	740,641.0	740,641.0
Geode Capital Management LLC	[2023-03-31]	DFND	722,045.0	722,045.0
BlackRock Inc.	[2023-03-31]	SOLE	654,893.0	657,023.0
Millennium Management LLC	[2023-03-31]	SOLE	344,684.0	344,684.0
Bridgeway Capital Management LLC	[2023-03-31]	SOLE	342,400.0	342,400.0
Element Capital Management LLC	[2023-03-31]	SOLE	331,135.0	331,135.0
State Street Corp	[2023-03-31]	DFND	207,153.0	207,153.0
Renaissance Technologies LLC	[2023-03-31]	SOLE	193,900.0	193,900.0
Morgan Stanley	[2023-03-31]	DFND	130,685.0	188,221.0
Susquehanna International Group LLP	[2023-03-31]	DFND	187,906.0	187,906.0
Jane Street Group LLC	[2023-03-31]	DFND	166,015.0	166,015.0
Prelude Capital Management LLC	[2023-03-31]	OTR	0.0	156,946.0
Citigroup Inc	[2023-03-31]	DFND	152,785.0	152,785.0
Prescott Group Capital Management LLC	[2023-03-31]	SOLE	126,970.0	126,970.0
HRT Financial LP	[2023-03-31]	DFND	117,480.0	117,480.0
Northern Trust Corp	[2023-03-31]	DFND	81.0	113,386.0
683 Capital Management LLC	[2023-03-31]	DFND	102,528.0	102,528.0
Brown Brothers Harriman & Co	[2023-03-31]	SOLE	100,000.0	100,000.0
Goldman Sachs Group Inc	[2023-03-31]	DFND	70,479.0	70,479.0
Barclays PLC	[2023-03-31]	SOLE	57,572.0	57,572.0
Qube Research & Technologies Ltd	[2023-03-31]	SOLE	0.0	46,977.0
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