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Type	Result	AUM ($BB)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
FMR LLC	[2023-03-31]	DFND	4,455,603.0	4,455,840.0
Price T Rowe Associates Inc /MD/	[2023-03-31]	SOLE	642,022.0	3,280,393.0
Column Group LLC	[2023-03-31]	SOLE	2,680,621.0	2,680,621.0
Perceptive Advisors LLC	[2023-03-31]	SOLE	2,291,608.0	2,291,608.0
BlackRock Inc.	[2023-03-31]	SOLE	2,136,495.0	2,171,704.0
State Street Corp	[2023-03-31]	DFND	1,578,969.0	1,639,046.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	1,516,523.0
Great Point Partners LLC	[2023-03-31]	DFND	0.0	1,355,151.0
OrbiMed Advisors LLC	[2023-03-31]	DFND	310,400.0	1,251,070.0
T. Rowe Price Investment Management, Inc.	[2023-03-31]	SOLE	486,062.0	1,231,783.0
Kingdon Capital Management LLC	[2023-03-31]	DFND	875,984.0	875,984.0
Morgan Stanley	[2023-03-31]	DFND	773,280.0	774,175.0
Redmile Group LLC	[2023-03-31]	SOLE	688,132.0	688,132.0
Cormorant Asset Management LP	[2023-03-31]	DFND	650,000.0	650,000.0
Deep Track Capital LP	[2023-03-31]	SOLE	571,233.0	571,233.0
Alphabet Inc.	[2023-03-31]	DFND	0.0	540,270.0
Geode Capital Management LLC	[2023-03-31]	DFND	528,555.0	528,555.0
Euclidean Capital LLC	[2023-03-31]	SOLE	432,900.0	432,900.0
Dimensional Fund Advisors LP	[2023-03-31]	SOLE	387,222.0	425,627.0
Citadel Advisors LLC	[2023-03-31]	DFND	374,445.0	374,445.0
Candriam SCA	[2023-03-31]	SOLE	330,399.0	330,399.0
Nuveen Asset Management LLC	[2023-03-31]	DFND	326,174.0	326,174.0
Point72 Asset Management LP	[2023-03-31]	DFND	0.0	325,700.0
Polar Asset Management Partners Inc	[2023-03-31]	SOLE	315,900.0	315,900.0
Boxer Capital, LLC	[2023-03-31]	OTR	300,000.0	300,000.0
Deutsche Bank AG	[2023-03-31]	DFND	5,432.0	295,894.0
Northern Trust Corp	[2023-03-31]	DFND	17,203.0	286,574.0
Fred Alger Management LLC	[2023-03-31]	SOLE	196,489.0	196,489.0
Charles Schwab Investment Management Inc	[2023-03-31]	SOLE	194,966.0	194,966.0
Susquehanna International Group LLP	[2023-03-31]	DFND	49,366.0	192,666.0
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