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Type	Result	AUM ($BB)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
EAM Investors LLC	[2023-03-31]	SOLE	0.0	6,675,132.0
OrbiMed Advisors LLC	[2023-03-31]	SOLE	5,559,528.0	5,559,528.0
RTW Investments LP	[2023-03-31]	SOLE	3,297,843.0	3,297,843.0
Cowen and Company LLC	[2023-03-31]	DFND	2,807,562.0	2,807,562.0
Millennium Management LLC	[2023-03-31]	SOLE	1,654,013.0	1,654,013.0
AIGH Capital Management LLC	[2023-03-31]	SOLE	1,565,149.0	1,565,149.0
Jefferies Financial Group Inc.	[2023-03-31]	DFND	1,316,000.0	1,316,000.0
5AM Venture Management LLC	[2023-03-31]	SOLE	1,200,647.0	1,200,647.0
Morgan Stanley	[2023-03-31]	DFND	1,037,067.0	1,167,561.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	994,184.0
Worth Venture Partners LLC	[2023-03-31]	OTR	0.0	510,648.0
AIGH Capital Management LLC	[2023-03-31]	OTR	0.0	510,648.0
King Luther Capital Management Corp	[2023-03-31]	SOLE	414,500.0	414,500.0
Susquehanna International Group LLP	[2023-03-31]	DFND	226,653.0	301,153.0
Two Sigma Investments LP	[2023-03-31]	SOLE	249,067.0	249,067.0
Kennedy Capital Management LLC	[2023-03-31]	SOLE	226,604.0	226,604.0
Bridgeway Capital Management LLC	[2023-03-31]	SOLE	192,200.0	192,200.0
Dimensional Fund Advisors LP	[2023-03-31]	SOLE	176,161.0	182,792.0
Renaissance Technologies LLC	[2023-03-31]	SOLE	137,712.0	180,986.0
Geode Capital Management LLC	[2023-03-31]	DFND	171,444.0	171,444.0
Laurion Capital Management LP	[2023-03-31]	SOLE	165,000.0	165,000.0
Jane Street Group LLC	[2023-03-31]	DFND	151,132.0	151,132.0
Two Sigma Advisers LP	[2023-03-31]	SOLE	120,600.0	139,700.0
Citadel Advisors LLC	[2023-03-31]	DFND	118,322.0	118,322.0
BlackRock Inc.	[2023-03-31]	SOLE	111,870.0	111,889.0
Goldman Sachs Group Inc	[2023-03-31]	DFND	109,132.0	109,132.0
Ameriprise Financial Inc	[2023-03-31]	DFND	0.0	96,597.0
Cubist Systematic Strategies, LLC	[2023-03-31]	DFND	0.0	91,453.0
ClariVest Asset Management LLC	[2023-03-31]	DFND	12,684.0	80,784.0
Squarepoint Ops LLC	[2023-03-31]	DFND	0.0	74,205.0
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