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Type	Result	AUM ($B)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
Vanguard Group Inc	[2025-06-30]	SOLE	0	6,528,778
BlackRock, Inc.	[2025-06-30]	SOLE	4,518,310	4,590,273
Marshall Wace LLP	[2025-06-30]	OTR	0	2,715,489
Caligan Partners LP	[2025-06-30]	SOLE	2,300,000	2,300,000
Geode Capital Management LLC	[2025-06-30]	DFND	2,200,428	2,200,428
Susquehanna International Group LLP	[2025-06-30]	OTR	1,489,672	1,746,772
Millennium Management LLC	[2025-06-30]	SOLE	1,127,225	1,127,225
State Street Corp	[2025-06-30]	DFND	982,672	1,112,705
Morgan Stanley	[2025-06-30]	DFND	545,698	1,084,478
Nuveen, LLC	[2025-06-30]	DFND	971,329	971,329
Northern Trust Corp	[2025-06-30]	DFND	683,893	683,893
Citadel Advisors LLC	[2025-06-30]	DFND	577,893	577,893
Goldman Sachs Group Inc	[2025-06-30]	DFND	465,506	465,506
GSA Capital Partners LLP	[2025-06-30]	SOLE	426,584	426,584
Diadema Partners LP	[2025-06-30]	SOLE	400,000	400,000
UBS Group AG	[2025-06-30]	DFND	309,687	309,687
Charles Schwab Investment Management Inc	[2025-06-30]	SOLE	294,106	294,106
Anson Funds Management LP	[2025-06-30]	DFND	275,000	275,000
Jane Street Group LLC	[2025-06-30]	DFND	46,682	255,482
Opus Capital Group LLC	[2025-06-30]	SOLE	240,000	240,000
Bank of New York Mellon Corp	[2025-06-30]	DFND	197,689	201,460
Two Sigma Investments LP	[2025-06-30]	SOLE	197,642	197,642
Barclays PLC	[2025-06-30]	SOLE	172,803	172,803
RhumbLine Advisers	[2025-06-30]	SOLE	39,430	135,789
Vanguard Group Inc	[2025-06-30]	DFND	0	126,021
XTX Topco Ltd	[2025-06-30]	DFND	76,900	76,900
LPL Financial LLC	[2025-06-30]	SOLE	0	67,560
JPMorgan Chase & Co	[2025-06-30]	DFND	16,792	61,892
MetLife Investment Management LLC	[2025-06-30]	DFND	58,844	58,844
Atom Investors LP	[2025-06-30]	SOLE	54,624	54,624
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