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Type	Result	AUM ($BB)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
RA Capital Management LP	[2023-03-31]	SOLE	0.0	22,748,880.0
FMR LLC	[2023-03-31]	DFND	7,813,013.0	7,813,036.0
Paradigm Biocapital Advisors LP	[2023-03-31]	SOLE	6,477,999.0	6,477,999.0
5AM Venture Management LLC	[2023-03-31]	SOLE	6,361,723.0	6,361,723.0
Laurion Capital Management LP	[2023-03-31]	SOLE	3,182,889.0	3,182,889.0
Artal Group S.A.	[2023-03-31]	DFND	2,792,805.0	2,792,805.0
Janus Henderson Group PLC	[2023-03-31]	DFND	0.0	1,764,877.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	1,553,558.0
Woodline Partners LP	[2023-03-31]	SOLE	1,111,123.0	1,111,123.0
Johnson & Johnson	[2023-03-31]	DFND	0.0	1,074,658.0
Baker Bros Advisors LP	[2023-03-31]	SOLE	633,848.0	633,848.0
Sarissa Capital Management LP	[2023-03-31]	SOLE	618,000.0	618,000.0
BlackRock Inc.	[2023-03-31]	SOLE	467,038.0	467,598.0
Millennium Management LLC	[2023-03-31]	SOLE	406,851.0	406,851.0
OUP Management Co LLC	[2023-03-31]	SOLE	374,714.0	374,714.0
Ensign Peak Advisors, Inc	[2023-03-31]	SOLE	346,087.0	346,087.0
Susquehanna International Group LLP	[2023-03-31]	DFND	330,963.0	330,963.0
Geode Capital Management LLC	[2023-03-31]	DFND	315,947.0	315,947.0
Thrivent Financial for Lutherans	[2023-03-31]	SOLE	181,539.0	181,539.0
Thrivent Financial for Lutherans	[2023-03-31]	DFND	133,670.0	133,670.0
Jane Street Group LLC	[2023-03-31]	DFND	129,124.0	129,124.0
JPMorgan Chase & Co	[2023-03-31]	DFND	127,939.0	127,939.0
Bridgeway Capital Management LLC	[2023-03-31]	SOLE	100,200.0	100,200.0
Goldman Sachs Group Inc	[2023-03-31]	DFND	93,279.0	93,279.0
Trustees of Columbia University in the City of New York	[2023-03-31]	SOLE	91,910.0	91,910.0
Citigroup Inc	[2023-03-31]	DFND	86,921.0	86,921.0
Morgan Stanley	[2023-03-31]	DFND	69,400.0	69,475.0
Northern Trust Corp	[2023-03-31]	DFND	48.0	67,636.0
LMR Partners LLP	[2023-03-31]	SOLE	60,298.0	60,298.0
Bollard Group LLC	[2023-03-31]	DFND	0.0	55,555.0
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