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Type	Result	AUM ($BB)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
Column Group LLC	[2023-03-31]	SOLE	9,400,290.0	9,400,290.0
Casdin Capital LLC	[2023-03-31]	SOLE	6,078,860.0	6,078,860.0
RA Capital Management LP	[2023-03-31]	SOLE	0.0	6,067,483.0
Price T Rowe Associates Inc /MD/	[2023-03-31]	SOLE	695,417.0	4,347,591.0
FMR LLC	[2023-03-31]	DFND	3,758,924.0	3,760,296.0
BlackRock Inc.	[2023-03-31]	SOLE	2,733,837.0	2,785,487.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	1,833,156.0
T. Rowe Price Investment Management, Inc.	[2023-03-31]	SOLE	491,178.0	1,564,525.0
Citadel Advisors LLC	[2023-03-31]	DFND	1,128,680.0	1,128,680.0
Regents of the University of California	[2023-03-31]	SOLE	960,957.0	960,957.0
Alphabet Inc.	[2023-03-31]	DFND	0.0	915,705.0
Euclidean Capital LLC	[2023-03-31]	SOLE	915,705.0	915,705.0
Geode Capital Management LLC	[2023-03-31]	DFND	790,790.0	790,790.0
Harvard Management Co Inc	[2023-03-31]	SOLE	673,270.0	673,270.0
State Street Corp	[2023-03-31]	DFND	462,500.0	496,800.0
StepStone Group LP	[2023-03-31]	DFND	462,163.0	462,163.0
Northern Trust Corp	[2023-03-31]	DFND	2,693.0	347,944.0
Trustees of Columbia University in the City of New York	[2023-03-31]	SOLE	40,218.0	291,828.0
Renaissance Technologies LLC	[2023-03-31]	SOLE	74,000.0	264,900.0
Bridgeway Capital Management LLC	[2023-03-31]	SOLE	181,900.0	181,900.0
Millennium Management LLC	[2023-03-31]	SOLE	160,608.0	160,608.0
Bank of New York Mellon Corp	[2023-03-31]	DFND	111,658.0	127,940.0
Two Sigma Investments LP	[2023-03-31]	SOLE	118,723.0	118,723.0
TD Asset Management Inc	[2023-03-31]	SOLE	114,930.0	114,930.0
Charles Schwab Investment Management Inc	[2023-03-31]	SOLE	113,033.0	113,033.0
GSA Capital Partners LLP	[2023-03-31]	SOLE	88,168.0	88,168.0
Susquehanna International Group LLP	[2023-03-31]	DFND	60,653.0	81,953.0
Morgan Stanley	[2023-03-31]	DFND	66,201.0	80,419.0
Nuveen Asset Management LLC	[2023-03-31]	DFND	74,559.0	74,559.0
Bank of America Corp /DE/	[2023-03-31]	DFND	4,591.0	70,108.0
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