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Type	Result	AUM ($BB)

Form 13F Holdings
Firm	Period	DFND	Voting	Shares
BlackRock Inc.	[2023-03-31]	SOLE	15,997,000.0	16,391,319.0
State Street Corp	[2023-03-31]	DFND	12,680,746.0	12,988,589.0
Perceptive Advisors LLC	[2023-03-31]	SOLE	11,900,255.0	11,900,255.0
Vanguard Group Inc	[2023-03-31]	SOLE	0.0	11,365,957.0
Nuveen Asset Management LLC	[2023-03-31]	DFND	9,032,313.0	9,032,313.0
D. E. Shaw & Co., Inc.	[2023-03-31]	DFND	8,200,090.0	8,211,573.0
Stonepine Capital Management LLC	[2023-03-31]	SOLE	7,661,690.0	7,661,690.0
Magnetar Financial LLC	[2023-03-31]	SOLE	7,074,722.0	7,074,722.0
Assenagon Asset Management S.A.	[2023-03-31]	SOLE	6,665,021.0	6,665,021.0
AWM Investment Company Inc	[2023-03-31]	SOLE	6,015,000.0	6,015,000.0
Millennium Management LLC	[2023-03-31]	SOLE	5,373,501.0	5,373,501.0
Invesco Ltd.	[2023-03-31]	DFND	0.0	4,826,616.0
Morgan Stanley	[2023-03-31]	DFND	3,618,990.0	4,466,102.0
Geode Capital Management LLC	[2023-03-31]	DFND	4,046,250.0	4,046,250.0
Velan Capital Investment Management LP	[2023-03-31]	SOLE	4,000,000.0	4,000,000.0
Adage Capital Partners GP LLC	[2023-03-31]	SOLE	3,624,558.0	3,624,558.0
Altium Capital Management LP	[2023-03-31]	SOLE	2,870,000.0	2,870,000.0
Two Sigma Investments LP	[2023-03-31]	SOLE	2,321,359.0	2,321,359.0
Fred Alger Management LLC	[2023-03-31]	SOLE	2,017,594.0	2,017,594.0
Two Sigma Advisers LP	[2023-03-31]	SOLE	1,765,200.0	2,008,600.0
Point72 Asset Management LP	[2023-03-31]	DFND	0.0	1,934,576.0
Bank of America Corp /DE/	[2023-03-31]	DFND	4,000.0	1,853,374.0
Northern Trust Corp	[2023-03-31]	DFND	44,898.0	1,846,165.0
Caligan Partners LP	[2023-03-31]	SOLE	1,662,613.0	1,662,613.0
Bridgeway Capital Management LLC	[2023-03-31]	SOLE	1,593,100.0	1,593,100.0
Boothbay Fund Management LLC	[2023-03-31]	OTR	0.0	1,234,504.0
Hudson Bay Capital Management LP	[2023-03-31]	SOLE	1,212,966.0	1,212,966.0
Oberweis Asset Management Inc	[2023-03-31]	SOLE	1,169,500.0	1,169,500.0
American Century Companies Inc	[2023-03-31]	DFND	1,044,237.0	1,115,270.0
Rafferty Asset Management LLC	[2023-03-31]	SOLE	981,231.0	981,231.0
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